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Range
Gray
Catbird breeds from the southern mainland of British Columbia, the
southern half of the Canadian Prairie provinces, southern Ontario,
southwestern Quebec, New Brunswick, Prince Edward Island, and Nova
Scotia south through Washington, eastern Oregon, Utah, and eastern
Texas, east to the Atlantic coast. Winters mainly on the coastal
fringe of the Gulf coast and eastern Mexico, south to Panama and
the Caribbean islands.
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Status
On the coast, rare to uncommon migrant
and summer visitant locally to the lower Fraser River valley, Squamish
River valley, Pemberton valley, and Bella Coola River valley; casual
elsewhere on the coast.
In the interior, an uncommon migrant
and summer visitant to the Southern Interior, Southern Interior
Mountains, and Central Interior ecoprovinces, except locally fairly
common in the Okanagan valley and in the vicinity of Williams Lake;
further north, a casual summer visitant in the SubBoreal Interior
and the Boreal Plains ecoprovinces; casual in winter in the Southern
Interior, accidental in the Southern Interior Mountains. |
Status
Change
No change. |
Nonbreeding
The Gray Catbird is sparsely but widely
distributed across the interior of southern British Columbia north
to Quesnel and Golden, and from the southern Rocky Mountains west
to Riske Creek in the Cariboo and Chilcotin areas and casually in
the Fraser Lowland on the south coast. In addition, there are several
occurrence records from outside the known breeding range, including
Sechelt, Comox, Bella Coola, Kitimat, Hazelton, Kleena Kleene, Bowron
Lake, and Fort St. John.
The Gray Catbird has been recorded from
sea level to 150 m on the coast and between 270 and 2,050 m in the
interior. It is most abundant in riparian thickets or in open deciduous
forests with an understorey of dense deciduous shrubs, small trees,
and scattered conifers. Preferred habitats are thickets along the
edges of streams, lakes, and ponds, fencelines, roadsides, ditches
and woodland edges, and shrub-choked gullies on sagebrush-grassland
slopes. Closed forest habitats are avoided. Vegetation in preferred
habitat often includes clustered wild rose, red-osier dogwood, saskatoon,
willows, water birch, black hawthorn, choke cherry, trembling aspen,
and Oregon-grape. Human-influenced habitats include shrubby gardens
and vine-tangled hedgerows along orchards and farm fields, often
with Himalayan blackberry on the coast. Gray Catbirds rarely venture
far from dense cover. They are heard more often than seen, and are
one of the few songbirds in British Columbia that frequently sing
at night.
Detection of non-singing birds is difficult;
thus, birds may be present, but silent, before territorial activities
are detected. Apparent spring migration begins relatively late,
normally from mid to late May, although in some years early migrants
arrive in mid to late April. In the Okanagan valley, 28 of 37 first-of-year
records occurred between 21 May and 1 June, with a mean and median
date of 23 May (Cannings et al. 1987). In the south, migration peaks
from late May to early June and ends abruptly. In the Williams Lake
area, at the northern edge of its range, the earliest Gray Catbird
arrivals are in mid-May, but few are seen until June. The autumn
migration occurs mainly in late August and early September. Information
on autumn departures is scarce, but by mid-September few catbirds
are reported. In some years, small numbers may remain into early
October in southern areas. Migratory movements are inconspicuous,
as catbirds are secretive. From evidence of large numbers of
catbirds killed by colliding with tall
structures, such as lighthouses and tall buildings, Root (1988)
concluded that it is a night migrant.
The only winter occurrences are individuals
at Nakusp, Irish Creek near Vernon, and Vaseux Lake.
The Gray Catbird has been recorded regularly
in British Columbia from 10 April to 15 October. |
Breeding
The Gray Catbird is a distinctive member
of the summer avifauna in the warm southern valleys of the interior
of British Columbia. Its highest breeding densities are in the Okanagan,
Similkameen, and South Thompson river valleys; it is slightly less
abundant in the valleys of the Kootenay and Columbia rivers. In
the interior, nesting has been confirmed north to Wasa, Revelstoke,
Horsefly Lake, and Williams Lake, and west to the Chilcotin River
in the vicinity of Riske Creek. The Gray Catbird also regularly
breeds in the riparian thickets along the Fraser River between Lytton
and Williams Lake, and in the valley of the San Jose River south
of Williams Lake. It is scarce in summer elsewhere in the central
portions of the interior, where breeding likely occurs north to
Quesnel.
On the coast, small numbers breed in
the Fraser Lowland, mainly in the Pitt Meadows area. The Gray Catbird
was reported as common in summer at Stuie in the Bella Coola River
valley (Laing 1942), although breeding there has not been confirmed.
The species reaches its highest numbers
in summer in the Okanagan valley of the Southern Interior. Breeding
Bird Surveys for both coastal and interior routes contain insufficient
data for analysis.
Breeding
habitat is similar to nonbreeding habitat, but thickets including
wild rose are preferred. General habitats for reported nests were
natural shrub thickets (80%) and human-influenced habitats (20%),
mainly gardens, orchards, vineyards, hedgerows, and uncultivated
land associated with farms. In the Okanagan valley, natural sites
are frequently in moist draws and riparian areas. Cannings et al.
(1987) state that only 2 of 93 nests were not in deciduous shrubs
or small trees; they were in dense garden firs.
The Gray Catbird breeds at elevations
below 100 m in the lower Fraser River valley and up to 1,100 m in
the interior.
Populations are small and local throughout
British Columbia, with pairs widely scattered in suitable habitat.
Surveys have revealed relatively dense concentrations (5 to 10 pairs)
at the north end of Okanagan Lake, the west side of Vaseux Lake,
the north end of Osoyoos Lake, and the east end of Williams Lake.
The Gray Catbird has been recorded breeding
in British Columbia between 9 May and 16 August. |
| Nests: 
Nests are usually built in dense tangles
of vegetation. Specific nest sites (n = 139) included wild rose
(47%), lilac bushes (25%), tall Oregon-grape (7%), red-osier dogwood
(4%), willows (4%), and 20 other species of shrubs and small trees.
The nest is a bulky assemblage of coarse material. The nest cup
is lined with fine grass, plant fibres, and other soft items. The
most consistently used nest materials were grass (84%), twigs (44%),
plant fibres (29%), rootless (21 %), and leaves (9%); a wide assortment
of other materials appeared in fewer nests. The heights above ground
for 156 nests ranged from 0.2 to 8 m, with 53% between 1 and 2 m.
Eggs:
Dates
for 137 clutches ranged from 10 May to 6 August, with 55% recorded
between 12 June and 2 July. Calculated dates indicate that nests
may contain eggs on 9 May. Egg laying begins about a month later
in the Cariboo and Chilcotin areas than in more southern areas.
Some pairs, in the Okanagan valley at least, are double-brooded.
The laying of first clutches peaks in early June, second clutches
about 9 July (Cannings et al.1987). It is not known what proportion
of second clutches followed the loss of a first clutch rather than
a successful first brood. In an Ontario study, only 3 of 22 pairs
that successfully raised 1 brood attempted a second, but pairs that
lost the first clutch routinely attempted a second nest (Scott et
al. 1988). The same study revealed a gradual decline in mean clutch
size as the season progressed.
Sizes of 122 clutches ranged from 1 to
6 eggs (1E-3, 2E-7, 3E-22, 4E-57, 5E-32, 6E-1), with 73% having
4 or 5 eggs. The mean clutch size in British Columbia was 3.8 eggs
(n = 124 clutches visited 2 or more times), which was almost
identical to the average number of 3.9 eggs per clutch laid in Ontario
between 13 May and 2 June (n = 53) (Scott et al. 1988).
Both Canadian studies revealed larger clutch sizes than those recorded
from Michigan, which averaged 3.3 eggs (n = 22 clutches
of 2 or more). The primary difference was in the large component
of five-egg clutches in British Columbia (25%), whereas no clutches
over four eggs were found in Michigan (Zimmerman 1963).
The incubation period in British Columbia
was 12 to 15 days (n = 11).
Nest
Success:
Of 59 nests found with eggs and followed
to a known fate, 37 produced at least 1 young, for a success rate
of 63%. The success rate in Michigan was 61% (n = 23) (Zimmerman
1963). In British Columbia, predation on eggs or young was suspected
in 17 cases, although confirmation was lacking. Reported predators
included squirrel or chipmunk (11 nests), domestic cat (3 nests),
shrew, gopher snake, and American Crow (1 nest each). Heavy rain
was thought to have caused the death of 1 brood.
Young: Dates for 108 broods ranged from
15 June to 16 August, with 54% recorded between 1 and 20 July. Sizes
of 94 broods ranged from 1 to 5 young (1Y-4, 2Y-12, 3Y-18, 4Y45,5Y-15),
with 67% having 3 or 4 young. The nestling period in British Columbia
was 12 to 15 days (n = 10). Family groups remained together
for a short time after the young left the nest.
Brown-headed Cowbird Parasitism: In British
Columbia, 5% of 183 nests found with eggs or young were parasitized
by the cowbird. None of eight coastal nests was parasitized. Most
of the parasitism occurred in the Southern Interior, but the overall
rate of 6% was higher than the 4% reported for the Okanagan valley
(Cannings et al.1987). In southern Ontario, Peck and James (1987)
reported a parasitism rate of 1.5% (n = 1,193), although
they were careful to mention that the percentage of attempted parasitism
was undoubtedly higher.
Three parasitized nests were closely
observed in British Columbia. In each, the cowbird egg(s) disappeared
from the nest within 2 days. A catbird from 1 nest that had received
2 cowbird eggs was seen carrying a cowbird egg in its beak, and
both eggs were missing from the nest a day after they were deposited.
The Gray Catbird is known to routinely eject cowbird eggs within
a day after they are laid (Nickel 1958; Friedmann 1963; Rothstein
1975; Scott 1977). The latter author suggested that the attentiveness
of the Gray Catbird, and its vigorous defence of the nest, made
it difficult for cowbirds to lay their eggs and remove 1 or more
eggs of the host. In British Columbia, there is a record of a cowbird
nestling together with a catbird nestling, but there are no records
in British Columbia of the catbird fledging a cowbird. Elsewhere
there are a few cases of successful parasitism (Woodward 1976; Lowther
1980). |
Remarks
The Gray Catbird was formerly known as
Catbird. Belles-Isles and Picman (1986c) report that the Gray Catbird
destroyed a variety of eggs of other species placed near the nest.
For additional information on breeding
biology and on distinguishing age and sex, see Bent (1948), Nickell
(1965), Johnson and Best (1980), and Suthers and Suthers (1990). |
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