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Range
Breeds
from northeastern British Columbia, northern Alberta, and central
Saskatchewan east to New Brunswick, south through southern British
Columbia to northern Baja California, and across the southern United
States from California to South Carolina. Winters across the United
States from California to Florida; occasionally further north. Resident
in southern California and much of Arizona.
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Status
On the coast, uncommon to fairly common
migrant and summer visitant on southeastern Vancouver Island in
the Georgia Depression Ecoprovince; rare to uncommon migrant and
summer visitant on the Fraser Lowland of the Georgia Depression,
accidental there in winter; casual on the Southern Mainland Coast
and accidental on the Northern Mainland Coast of the Coast and Mountains
Ecoprovince.
In the interior, fairly common migrant
and summer visitant in the Southern Interior Ecoprovince; uncommon
to fairly common in the southern portions of the Southern Interior
Mountains and Boreal Plains ecoprovinces; very rare in the southern
Central Interior, the Sub-Boreal Interior, and Taiga Plains ecoprovinces.
Breeds. |
Status
Change
No change. |
Nonbreeding
The
House Wren has a widespread distribution during spring and summer
across extreme southern British Columbia from the south and east
coast of Vancouver Island north to Campbell River, east through
the lower Fraser River valley to Manning Park and north to the Nicola
valley; through the Okanagan valley, north to Adams Lake and Clearwater
(Dickinson 1953); and through southern portions of the west and
east Kootenay. In the Central Interior, there are fewer than 20
records, all from the area between 100 Mile House, Williams Lake,
and Riske Creek. A specimen from the Kispiox valley is the only
record from the Northern Main land Coast of the Coast and Mountains
Ecoprovince (Swarth 1924; Brooks and Swarth 1925). In the Sub-Boreal
Interior, there are 4 known localities where this wren has occurred:
Chetwynd, Prince George (Godfrey 1986), Quesnel, and Smithers. The
House Wren also occurs regularly in the Peace Lowland, and locally
but fairly regularly in the Fort Nelson area.
On the coast, it occurs mainly at elevations
from sea level to about 300 m, but there is 1 record at about 1,400
m. In the interior, it has been found up to 1,200 m.
The House Wren is a bird of drier habitats.
In the southern interior, it is most abundant in deciduous groves
within ponderosa pine forests (Cannings et al. 1987), but also occurs
in relatively open stands of trembling aspen, as it does in the
northeast. On Vancouver Island and the Gulf Islands, it is most
numerous along the edges of dry forests featuring arbutus, Carry
oak, and Douglas-fir, particularly in brushy areas at the forest
edge or along sun-baked rocky bluffs that border the sea. Specific
habitats include hedgerows, bramble tangles, thickets of Scotch
broom and gorse, open brushy areas along roads and powerline rights-of-way,
and thickets around ponds and seashore. Throughout its range, it
occurs frequently in orchards, gardens, and abandoned homesteads.
On the coast, spring migration begins
slowly in early April, rarely in late March, and peaks in May. East
of the coastal mountain ranges, the House Wren arrives after the
first week of April and numbers continue to increase into May and
June. In the Peace Lowland, spring migrants arrive in May and numbers
peak in late May and early June. The northeastern population is
continuous with that of western Alberta (Semenchuk 1992), and reaches
that area via a migration route east of the Rocky Mountains. West
of the Rocky Mountains, the House Wren migrates through intermountain
valleys.
In the Peace Lowland, the southward migration
occurs mainly in August, and most birds have gone by mid-September.
In the Southern Interior, the autumn movement can be detected as
early as late July but most birds leave in August, with few reported
in September and only casual occurrences thereafter. On the coast,
the autumn migration is more protracted, occurring in August and
September, with only a few individuals left in October and November.
There is only 1 convincing late winter
record for the House Wren in British Columbia: a specimen from Chilliwack
(see also REMARKS). Root (1988) states that the House Wren winters
along the Pacific coast mostly where the average January minimum
temperature does not drop below -1°C. Our data suggest that
even the relatively mild winters on the coast of British Columbia
are not suitable for this wren.
On the coast, the House Wren has been
recorded regularly from 27 March to 28 October; in the interior,
it has been recorded from 13 April to 6 October. |
Breeding
The
House Wren breeds throughout most of its summer range, from southeastern
Vancouver Island across the extreme southern portions of the province
to the foothills of the Rocky Mountains. A disjunct population breeds
in the Peace River region and at Fort Nelson. There are many nesting
records from the valleys of the Thompson, Okanagan, and Similkameen
rivers; in the Columbia River valley near Revelstoke, and the southern
west Kootenay; and near Cranbrook. The northernmost breeding records
west of the Rocky Mountains are near 100 Mile House, Williams Lake,
Quesnel, and Prince George.
The highest numbers in summer occur in
the Southern Interior, especially in the Okanagan valley. House
Wrens are also locally common breeders on southeastern Vancouver
Island, the Gulf Islands, and the Peace Lowland. On the lower Fraser
River delta, within a few kilometres of the sea, there are many
records of House Wrens in spring and summer, but few nesting records.
Beyond the centre of its range, the House
Wren may breed irregularly. In the Prince George area, this wren
seemed to disappear in the 1970s, but had returned to breed by 1993
(J. Bowling pers. comm.). In the Arrow Lakes valley north of Castlegar,
House Wrens were not recorded from 1975 to 1990, but several singing
males were found in the early 1990s (G.S. Davidson pers. comm.).
An analysis of Breeding Bird Surveys
for the period 196S through 1993 shows that the number of birds
on interior routes has increased at an average annual rate of 4%.
Data for coastal routes were insufficient for analysis.
In the interior, breeding has been recorded
at elevations from 270 m to about 1,150 m; on the coast, from near
sea level to 390 m. Most House Wren nests were reported from human-altered
habitats (79%; n = 832) and forests (12%), but this summary
is biased by the predominance of nests studied at 1 rural residence
in the Okanagan valley. Deciduous and mixed woodlands were preferred
to coniferous forest.
Specific nesting habitats included brushy
backyards or farmyards (59G/70; n = S12), pasture and open
rangeland (13%), woodlands (11%), orchards (8%), and riparian situations
(6%). Other sites included overgrown fields with brambles, fence
rows, and railway rights-of-way.
In the interior, the House Wren has been
recorded breeding from 26 April (calculated) to 28 August; on the
coast, it has been recorded from 25 April (calculated) to 12 August. |
| Nests: 
Most
nests were found in nest boxes (84%; n = 942), cavities such
as abandoned woodpecker nests or natural cavities left by the rotting
of branches (13%), and crevices (2%). House Wrens often used nest
boxes set out for bluebirds or swallows. Specific sites for non-nest
box nests (n = 130) included deciduous trees (58%), stumps
(16%), coniferous trees (13%), and buildings (12%). The heights
for 139 nests in natural cavities ranged from 0.9 to 15 m, with
55% between 1.8 and 4.5 m.
Nest materials included mainly twigs
(99%; n = 813), feathers (65%), grass (24%), hair (3%), moss,
and occasionally spider webs, rootlets, mud, rope, and leaves. Nests
varied in size, as materials usually filled the available space
in cavities. The lining of the nest cup usually included fine grass,
animal hair, feathers, plant fibres, leaves, plant stalks, pine
needles, and other fine materials.
Eggs:
Dates for 707 clutches ranged from 28
April to 24 August, with 52% recorded between 1 June and 29 June.
Calculated dates suggest that nests can have eggs as early as 25
April. Sizes of 684 clutches ranged from 1 to 9 eggs (1E-17, 2E12,
3E-25, 4E-53, 5E-123, 6E-167, 7E-220, SE-59, 9E-S), with 57% having
6 or 7 eggs. In 3 nests, the incubation period was between 14 and
15 days. In Ontario, Peck and James (1987) give a range of 10 to
19 days, with an average of 12 to 14 days.
Cannings et al. (1987) note that clutch
size in the Okanagan valley declines through the summer; 146 clutches
started before 15 June had a mean clutch size of 6.4 eggs, while
71 clutches initiated later than that had a mean clutch size of
5.2 eggs. About half of the nesting females in the Okanagan valley
attempt to produce 2 broods each breeding season.
Nest
Success:
Of 535 nests found with eggs and followed
to a known fate, 449 produced at least 1 fledgling, for a success
rate of 84%; coastal nest success was 100% (n = 2); interior
nest success was 84% (n = 533).
Finch (1989) showed that the nesting
success of House Wrens was greater when nests were placed in open
habitats. Belles-Isles and Picman (1986b) demonstrated the same
results and suggested that greater nesting success in sparser vegetation
resulted from a more effective defence of nests against other marauding
House Wrens in such sites.
Young: Dates for 683 broods ranged from
15 May to 28 August, with 51% recorded between 18 June and 17 July.
Sizes of 598 broods ranged from 1 to 11 young (1Y-9, 2Y-7, 3Y-25,
4Y-51, 5Y-132, 6Y-153, 7Y-175, SY-43,9Y-3), with 55% having 6 or
7 young. In 13 nests, the nestling period ranged from 14 to 22 days,
with a median of 16 days.
Brown-headed Cowbird Parasitism: Cowbird
parasitism was not found in British Columbia in 802 nests recorded
with eggs or young. In Ohio, Murphy (1984) reported 2 cases of cowbird
parasitism in a sample of over 900 House Wren nests. In Ontario,
Peck and James (1987) found 4 nests parasitized in a sample of 1,120
nests. Friedmann and Kiff (1985) list only 9 records for North America.
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Remarks
Three subspecies of House Wren occur
in North America; the one occurring in British Columbia is T.
a. parkmanii (Godfrey 1986).
All winter records of the House Wren
must be carefully documented. We have a number of winter reports
of this wren on file, including published Christmas Bird Counts
from both the coast and the interior - Campbell River, Comox, Ladner,
Nanaimo (including the all-time Canadian high count of 4 birds on
30 Dec 1979 [Anderson l9S0]), Vancouver, Victoria, and Shuswap
Lake Provincial Park (Appendix 2). All lack adequate details, and
we have excluded them from this account. In addition, 10 House Wrens
recorded on the Telkwa High Road Breeding Bird Survey on 23 June
1974 are also without documentation and have not been considered
here. The House Wren is notorious as a nest predator. Territorial
House Wrens are known to destroy nests of other species and other
House Wrens, whether or not they are within its nesting territory.
The behaviour includes removing the soft nest lining, killing small
nestlings, and pecking holes in eggs
(Hill 1869; Sherman 1925; Creaser 1925;
Kendeigh 1941; Belles Isles and Picman 1986a). Experiments indicate
that this behaviour has the effect of making preferred nesting sites
available for their own use (Pribil and Picman 1991).Kennedy and
White (1992) discuss nest box selection by the House Wren, and conclude
that boxes with wide, slot shaped entrances were more likely to
contain a nest with eggs, apparently because the male could more
readily carry twigs through slots than holes. McCabe (1961, 1965)
and Lumsden (1986) provide additional information on the use and
selection of nest boxes by House Wrens. |
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