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Range
Breeds
from central Alaska and Yukon, southwestern Mackenzie, northern
British Columbia, southwestern Alberta, and Saskatchewan south through
the western United States to Baja California and southern Mexico.
Winters from central coastal California to southern Mexico and northern
Central America.
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Status
Status is complex. On the coast, common
to very abundant spring migrant on southeastern Vancouver Island
and the entire lower Fraser River valley in the Georgia Depression
Ecoprovince. Fairly common to very common summer visitant and autumn
migrant there; occasionally abundant in autumn and very rare in
early winter. Uncommon to locally common migrant and summer visitant
to Western Vancouver Island and the southern and northern portions
of the Coast and Mountains Ecoprovince; very rare transient on the
Queen Charlotte Islands.
In the interior, fairly common to very
common (occasionally abundant) spring migrant and summer visitant
to the south-central portions of the province, including the Southern
Interior and Southern Interior Mountains ecoprovinces; common to
abundant (occasionally very abundant) autumn migrant; casual in
winter there. Northward, throughout the rest of the province, uncommon
to locally common migrant and summer visitant except in the extreme
northeastern corner, where it is a casual spring transient in the
Taiga Plains Ecoprovince.
Breeds. |
Status
Change
No change. |
Nonbreeding
The
Violet-green Swallow is widely distributed across southern British
Columbia except for the west coast of Vancouver Island and the central
mainland coast, where it is localized and scarce in some years.
It is less numerous through the Cariboo and Chilcotin areas, and
its distribution is further reduced in the northern parts of the
province. Unlike the Tree Swallow, the Violet-green Swallow seldom
appears on the Queen Charlotte Islands or in the Taiga Plains, and
then only during migration.
The Violet-green Swallow occurs from
near sea level to 1,400 m elevation. Migrating flocks closely follow
rivers, lakes, and valley bottoms as they move northward during
the spring migration before dispersing into breeding areas. Both
on the coast and in the interior, it seems to favour lowland areas
with open terrain and concentrations of flying insects, on which
it feeds. It is usually found over freshwater habitats, including
large and small lakes, rivers, marshes, bogs, wet meadows, and beaver
ponds; adjacent forest is also used. This swallow forages from low
over water or ground surfaces to heights of 100 m or more above
a forest canopy. The Violet-green Swallow occurs less frequently
along the coast except over estuaries, where it can be found in
brackish meadows, mudflats, fields, and grassy beaches. Rural and
urban areas are also used, particularly agricultural pastures and
well-treed suburbs. In the early spring, sunny cliff faces may also
provide important foraging sites.
The timing of the spring migration varies
widely between years. Violet-green Swallows are among the earliest
returning swallows and are particularly dependent on weather conditions,
which determine the timing of insect hatches. In exceptionally mild
winters, the occasional bird can arrive at the south coast in early
February, and in some years the first flocks arrive in late February.
Normally, however, large numbers of Violet-green Swallows appear
suddenly on the Fraser River delta and southeastern Vancouver Island
in March, and increase through April. These are birds in passage.
Their numbers decline abruptly in May and drop further in June and
July. A similar trend is evident from other areas along the coast.
The coastal movement peaks about 2 weeks ahead of migration through
the southern interior.
In southern portions of the interior,
birds may arrive in late February and early March; their numbers
peak in April and then decline. Spring migration through the Southern
Interior and Southern Interior Mountains is smaller and less spectacular
than the coastal movement. In the Cariboo and Chilcotin areas of
the Central Interior, the peak movement occurs in the second and
third weeks of April. Few birds move through the Sub-Boreal Interior.
In the northern interior, early arrivals occur in mid to late April,
becoming most numerous in early May.
In spring, the Violet-green Swallow usually
migrates in flocks of 20 to 200 birds, although flocks of thousands
can occur. Of 86 aggregations with more than 200 birds, 84% occurred
in the Georgia Depression, 2% in the Southern Interior, 13% in the
Southern Interior Mountains, and 1% elsewhere. During peak spring
movements, large numbers may accumulate in good foraging areas.
For example, all records of more than 1,000 birds are from lakes,
all in the extreme south. The 5 maximum counts (2,500 to 9,000 birds)
were all recorded in April or the first week of May, at Beaver and
Elk lakes, north of Victoria. The highest total count of Violet-green
Swallows in the province is 11,000 birds, tallied near Victoria
at Elk, Beaver, and Prospect lakes on 15 April 1970 (Tatum 1971).
Southeastern Vancouver Island has the largest spring movements in
the province.
Post-breeding flocks congregate over
larger lakes, rivers, and marshes in July and August before moving
southward. During midsummer, mixed flocks of swallows, numbering
hundreds of birds, are common sights over interior lakes.
The autumn migration is more protracted
than the spring movement and involves smaller flocks of Violet-green
Swallows. Peaks vary between years, but generally their movements
begin earlier than those of other swallows. Numbers recorded on
the southern migration also suggest different behaviour than during
the spring movement. On the coast, where there is such a dramatic
northern movement in spring, swallows depart almost unnoticed in
autumn, and most have left by the first week of October. In the
southern portions of the interior, however, almost the reverse is
true. Of 24 autumn flocks of 200 birds or more, only 12% were from
the south coast while 88% were from the southern portions of the
interior.
In the northern and central regions,
migration begins shortly after nesting ends. The highest numbers
occur in July, and most birds have gone by early to mid-August.
In the Okanagan, Kootenay, and Columbia valleys, the highest numbers
occur in August and September, and most birds have gone by the first
week of October. In the Okanagan valley, the autumn migration peaks
in early September (Cannings et al. 1987). Almost all Violet-green
Swallows have left the province by the end of September.
The pattern of records suggests different
routes for spring and autumn migration. In spring, most Violet-green
Swallows move northward along the coast, taking advantage of the
milder weather and available food. The small number of birds that
migrate in spring up the valleys of the southern portions of the
interior suggests that the nesting population in the Central Interior
and beyond is derived from the large coastal concentrations that
move into the interior along major west-east river valleys. Following
the breeding season, coastal birds leave shortly after their young
fledge and no noticeable concentrations occur. For birds in the
interior, there is no advantage in returning to the coast. Swarms
of late-summer chironomids and other flying insects create an abundant
supply of food over lakes and ponds. Observations of large numbers
indicate that the swallows leave through the Okanagan, Creston,
and other southern valleys. Mountain passes and montane areas (e.g.,
Manning Park) may also be important autumn migration corridors,
but their use has not been well documented.
Although migrating swallows spend much
of their time on the wing, perching and roosting sites are important
habitat components. Frequently, they use snags, leafless trees or
branches, and roots protruding from eroding river banks. In rural
and urban areas, telephone lines, buildings, wire fences, and bridges
also provide perching sites.
On the coast, the Violet-green Swallow
has been reported every month of the year, although it occurs in
substantial numbers only from late February to late September. In
the interior, it has been reported from 19 February to 8 November. |
Breeding
The
Violet-green Swallow breeds throughout most of its range in the
province. It is scarce in the Peace Lowland and in the Northern
Boreal Mountains. The northernmost breeding records are from the
Tatshenshini River, Atlin, Fireside, and Sulphur Creek, along the
northern boundary of the province. There is a vast area west of
the Rocky Mountains and north of the latitude of Fort St. John where
there have been few observers and thus few nest records. For example,
although Swarth (1922) found them abundant at Telegraph Creek, there
are no other nesting records in the Stikine River drainage. There
are just 3 nesting records on the coast north of Vancouver Island:
Owikeno Lake, Kitsumkalum Lake, and Alice Arm. The centre of breeding
abundance for the Violet-green Swallow is in the Georgia Depression,
Southern Interior, and Southern Interior Mountains, where it has
adapted well to human-influenced habitats. Breeding has not been
reported from the Queen Charlotte Islands or the Taiga Plains.
This species reaches its highest numbers
on southeastern Vancouver Island and in the lower Fraser River valley
of the Georgia Depression. An analysis of Breeding Bird Surveys
for the period 1968 through 1993 could not detect a net change in
numbers on either coastal or interior routes. Both Robbins et al.
(1986) for North America and Erskine et al. (1992) for southern
British Columbia also note stable Violet-green Swallow populations.
On the coast, the Violet-green Swallow
has been found breeding from near sea level to 390 m elevation;
in the interior, it breeds from 300 to 1,400 m elevation. It seems
to thrive in close association with humans (Weber 1972; Allen et
al. 1977). Over 75% of 847 nests were from human-associated habitats.
Most nests (62%; n = 805) were found in backyard gardens and residential
buildings, roadside habitats, meadows, or shorelines (lakes, streams,
marshes, seashores). Only 6% were reported from cliffs, which suggests
a bias due to observer distribution. Other habitats included orchards,
pasture and other farmland, parks, golf courses, and campgrounds.
The Violet-green Swallow has been recorded
breeding in the province from 1 April (calculated) to 24 August. |
| Nests: 
Although
the Violet-green Swallow nests solitarily, it also nests in substantial
colonies, particularly on cliffs. Colony size ranged from 3 to 40
pairs (n = 16), with 8 between 5 and 15 pairs. Thirteen of the colonies
were found in the interior.
The Violet-green Swallow has adapted
well to nesting in human-made habitats such as nest boxes, the crannies
of houses, garages, sheds, barns, and other buildings, and in posts
and poles, stone and brick walls, and bridges. Natural nest sites
include rock cliffs, snags, and cavities in living deciduous and
coniferous trees. Along the Fraser River canyon, near Williams Lake,
it nests all along the cliffs, but has never used any of the more
than 1,000 boxes on the adjacent plateau (A. Roberts pers. comm.).
Violet-green Swallows seem to prefer boxes on buildings rather than
those on poles and posts. In suburban environments, the House Sparrow
frequently displaces the Violet-green Swallow from nest sites (Erskine
1979).
In the Cariboo and Chilcotin areas, about
half of all nests were found in natural situations. Further north,
in the SubBoreal Interior and Boreal Plains, natural sites such
as crevices in cliff faces and cavities in a variety of tree species
were the preferred nesting sites.
The adaptability of the Violet-green
Swallow is indicated by the diversity of nest sites reported; any
darkened cavity with an adequate entrance hole or crevice seems
suitable. The species has been reported nesting inside the walls
of buildings and on fiberglass bats in attics where they have gained
access through knotholes or ventilation louvres, under tiles on
the ridges of house roofs, between logs in log buildings, under
roof shingles and flashing, in chimneys, in a trailer hitch of a
semi-trailer truck, under a plywood sheet in a lumberyard, in plumbing
vent pipes of buildings, and in the blasting holes of roadside rock-cuts.
Sometimes a human-made site can prove to be a death trap for the
birds. In the mid-1970s, dismantling of a wooden shed on the Alaksen
National Wildlife Area, south of Vancouver, revealed nearly 100
swallow skeletons between the studs of a wall (R. McKelvey pers.
comm.). The knothole through which they had entered was about 1.2
m up the wall, and the remains filled the space between the studs
to a depth of about 0.8 m. After having gained access, the swallows,
for some reason, could not return to the hole to exit.
Reported nests were loosely constructed,
the amount of material governed by the size of the cavity occupied.
The nest was usually a collection of grasses with a small cup formed
in the centre or in 1 corner of the cavity; the cup was formed with
finer grasses and lined generously with feathers. White feathers
seemed to be preferred. Most nests (94%; n = 515) were made of grasses
and were feather-lined (80%). One nest was made entirely of feathers.
Other materials included plant fibres, rootlets or twigs (6%), mud
(5%), hair (3%), conifer needles, wood chips, plant down, and human-made
materials such as fiberglass insulation, string, rope, and paper.
Nest sites are reused in successive years.
The heights of 460 nests in nest boxes
ranged from 0.3 to 21 m, with 54% between 2.4 and 4.2 m; 344 nests
where nest boxes were not used ranged from ground level to 78 m,
with 62% between 2.4 and 6.0 m. Many of the known cliff sites have
not been included in the samples.
Eggs:
Dates for 302 clutches ranged from 7
April to 21 July, with 51% between 29 May and 16 June. Calculated
dates indicate that eggs can occur as early as 1 April and as late
as 1 August. Clutch size ranged from 1 to 8 eggs (1E-13, 2E-18,
3E-20, 4E-92, 5E-117, 6E-37, 7E-4, 8E-1), with 69% having 4 or 5
eggs. The 8-egg clutch may have been the product of 2 females. The
incubation period in British Columbia ranged from 14 to 17 days
(n = 16), with a median of 15 days, similar to the 14 or 15 days
reported by Turner and Rose (1989) and Brown et al. (1992).
Nest
Success:
Of 188 nests found with eggs and followed
to a known fate, 139 produced at least 1 fledgling, for a success
rate of 74%. Coastal nest success was 76% (n = 82); interior success
was 74% (n = 106). Weather and parasitism by the larvae of Protocalliphora
sp. can have significant negative impact on the breeding success
of the Violet-green Swallow (see also Tree Swallow and Barn Swallow).
Other causes of egg or young mortality in British Columbia included
predation by House Sparrows, chipmunks, squirrels, crows, deer mice,
Cooper's Hawks, and American Kestrels; and nest usurpation by European
Starlings, Crested Mynas, and House Sparrows. There are 2 reports
of birds laying second clutches after the first brood was successfully
raised; there are a number of reports of renesting following the
loss of eggs.
Young: Dates for 416 broods ranged from
15 April to 24 August, with 53% between 20 June and 9 July. Sizes
of 353 broods ranged from 1 to 7 young (1Y-20, 2Y-50, 3Y-7, 4Y-142,
5Y-105, 6Y-26, 7Y-3), with 59% having 4 or 5 young and 76% with
3 to 5 young. The nestling period in British Columbia ranged from
23 to 27 days (n = 14), with a median of 25 days. Edson (1943) found
a nestling period of 23 days.
Brown-headed Cowbird Parasitism: In British
Columbia, cowbird parasitism was not found in 800 nests recorded
with eggs or young; the Violet-green Swallow has not been found
as a cowbird host elsewhere in North America either (Friedmann et
al. 1977; Friedmann and Kiff 1985). |
Remarks
Godfrey (1986) shows the breeding distribution
of the Violet-green Swallow to include the Queen Charlotte Islands
and the Taiga Plains Ecoprovince in extreme northeastern British
Columbia; however, we could find no evidence that this swallow breeds
in those areas.
British Columbia is the centre of Violet-green
Swallow breeding abundance in Canada. Yet, as Brown et al. (1992)
note, "Despite an extensive distribution, less is known about the
Violet-green Swallow than nearly any other North American Swallow."
Their review could find no recent studies of the species' breeding
biology or behaviour and was based primarily on reports from the
1940s and 1950s. Interested researchers in British Columbia have
the opportunity to add to our knowledge of this species.
Erskine (1979) notes that the Violet-green
Swallow's ability to nest in remote cliffs and tree cavities and,
at the same time, adapt well to human habitation may have saved
it from harmful human impact. It readily takes to nest boxes, and
once established will return year after year; an adult female banded
at a nest box on 30 June 1983 at the Qualicum National Wildlife
Area, Vancouver Island, was recaptured in successive years in the
same or a nearby nest box until 18 June 1989.
Since the Violet-green Swallow is a secondary
cavity-nester, its nesting density may be limited by a scarcity
of suitable nest sites in regions lacking rock cliffs (Brawn and
Balda 1988).
See Brown et al. (1992) for a summary
of the life history of the Violet-green Swallow in North America. |
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